They are trees or shrubs; with terete to sub-square distal twigs, variously hairy to glabrous. Leaves mostly 5-25 cm, chartaceous to subcoriaceous, whole to toothed, undulate-denticulate or serrulate. Flowers 4-12-mere and typically polystane, in terminal or subterminal panicles (rarely pseudo-axillary), multifloral, bract. Hypanthic domed, campanulate, or hemispherical, glabrous inside and typically thick-walled; calyx calyptriform, circumcisile at or near the base at anthesis.